Based on Dr. Norton's articles, below is an outline of the angelfish phenotype timeline. New genes
have emerged recently and are still being studied, therefore... they are not included in this list.
Dark (D) Dominant
1953 Europe, blacklace; 1954 Julianna & Fred Ludwig all black angelfish Detroit Michigan
Innes July 1955
Wolfsheimer July 1955
Halfblack (h) Recessive
Appeared in the hobby approximately 1955.
There is no known documentation of it origin (Axelrod, 1985).
Stripeless (s) Recessive
1965 Conles blushing-named by Lester Boisvert, produced by Vorderwinkler
Marble (M) Dominant
1969 Marble-Charles Ash
Naja gold (sometimes called Wingate gold) (Lost)
1970 Carl Naja (The first gold to enter the hobby and is now obsolete.)
Smokey (S) Dominant
1971 Smokey-Charles Ash
Hong Kong gold (Lost)
1971 Anon. Peter Wong (This
gene may also be obsolete.)
Zebra (Z) Dominant
1971 Origin is unknown. Entered the trade in the 70's.
New gold (g) Recessive
1973 Imported into the US from Singapore 1974 Earl Wellwood, a Canadian 1975 Ross Socolof
Gold marble (Gm) Dominant
Apparently it originated in the Far East. Timeframe is unknown.
Streaked (St) Dominant
First encountered sometime in the early 80's. No known
documentation of it's origin. Dr. Norton first obtained her streaked angelfish from
Tim Anderson. Year is unknown.
Pearlscale (p) Recessive
1987 by Kamphol Udomritthiruj (Seattle). 1989 Tim Anderson, a tropical fish wholesaler and angelfish breeder in
Eagle Grove, Iowa, imported some pearly angelfish from Bangkok early in
1989. The supplier had changed it's name to "Pearly" however
many breeders still use its original name today.
